拉斯克基础医学研究奖、沃尔夫医学奖、托马斯·亨特·摩尔根奖章、哈维奖、盖尔德纳国际奖、国际生物学奖、路易莎·格罗斯·霍维茨奖、NAS Award in the Neurosciences、罗森斯特尔奖、Karl Spencer Lashley Award、Bristol-Myers Squibb Award for Distinguished Achievement in Neuroscience Research、Robert J. and Claire Pasarow Foundation Award for Distinguished Contributions to Neuropsychiatric Research、Crafoord Prize in Biosciences、Bower Award and Prize for Achievement in Science、英国皇家学会外籍会员、Ralph W. Gerard Prize、honorary doctor of Purdue University、Mendel Medal、Howard Taylor Ricketts Prize