哈里猪笼草

哈里猪笼草
	产于基纳巴卢山登顶步道沿途的哈里猪笼草的下位笼
保护状况
	瀕危
科学分类
界：	植物界 Plantae
演化支：	维管植物 Tracheophyta
演化支：	被子植物 Angiosperms
演化支：	真双子叶植物 Eudicots
目：	石竹目 Caryophyllales
科：	猪笼草科 Nepenthaceae
属：	猪笼草属 Nepenthes
种：	哈里猪笼草 N. × harryana
二名法
	Nepenthes × harryana; Burb. (1882)

哈里猪笼草（学名：Nepenthes × harryana）是由爱德华猪笼草（N. edwardsiana）和长毛猪笼草（N. villosa）杂交得到的自然杂交种。^[3]其亲本近缘关系非常密切，其形态特征为亲本的中间型，较难进行区分。^[4]^[5]其得名于园艺公司维奇苗圃的负责人哈里·维奇。

植物学史

1882年，弗雷德里克·威廉·伯比奇将其作为一个物种进行了最初的描述。^[2]他写道：^{[注 1]}^[2]

“

除此之外，我还发现了一种为长毛猪笼草和爱德华猪笼草的中间型的物种附生于木麻黄上。我相信这是一个未命名的物种；如果是，我想将其称为哈里猪笼草。对爱德华猪笼草的枯萎捕虫笼进行观察可发现，其上五分之四为纸质，而下部为坚硬的革质；长毛猪笼草除了唇下约半英寸外，几乎全为坚硬的革质；哈里猪笼草约有三分之一的捕虫笼是革质的，而唇下的三分之二为柔软的纸质。在我的草图中可看到，这三者的笼口边缘与其他猪笼草明显不同。

……

约瑟夫·胡克先生在《Linn. Trans.》第二十二卷中认为长毛猪笼草与爱德华猪笼草可能是同一个物种。但事实并非如此，他们在山上分布的区域距离很远，且生长习性及颜色都不一样。而哈里猪笼草显然是一个杂交种，其具有双亲的性状。

”

1908年，约翰·缪尔黑德·麦克法兰在他的专著《猪笼草科》中跟随了弗雷德里克·威廉·伯比奇的杂交种假设并进行了描述。^[6]B·H·丹瑟在其1928年的专著《荷属东印度群岛的猪笼草科植物》中也同麦克法兰一样，认为哈里猪笼草是一个杂交种，并认为其可能是爱德华猪笼草与长毛猪笼草的一个变型的自然杂交种。但在其最后的解释中，他将这两个类群都归入了长毛猪笼草。^[7]

注释

^ 弗雷德里克·威廉·伯比奇对哈里猪笼草的叙述原文：^[2]

“

Apart from these I found an intermediate between N. villosa and N. Edwardsiana, also epiphytic on Casuarina. This is, I believe, unnamed ; if so, I should like it to be called Nepenthes Harryana. Now, if a dried pitcher of N. Edwardsiana be examined, the upper four-fifths of it will be seen to be membranous, the lower part leathery and hard ; in N. villosa nearly all is hard and leathery except about half-an-inch below the hardened rim of the urns ; in N. Harryana about one-third is hard, and two-thirds soft or membranous below the rim. The edge of the pitcher mouths in these three kinds is quite distinct from those of all others, as shown in my sketches.

[...]

Sir Joseph Hooker, in Linn. Trans., vol. xxii., suggested that N. villosa and N. Edwardsiana might be forms of the same species. This is not so, however, they are quite distinct in zone on the mountain, and in habit of growth also, and in colour. Then N. Harryana is a hybrid no doubt, which shows they have distinct sexual characters.

”

参考文献

^ von Arx, B., J. Schlauer & M. Groves 2001. CITES Carnivorous Plant Checklist.PDF The Cromwell Press, United Kingdom.
^ ^2.0 ^2.1 ^2.2 ^2.3 Burbidge, F.W. 1882. Notes on the new Nepenthes. （页面存档备份，存于互联网档案馆） The Gardeners' Chronicle, new series, 17(420): 56.
^ McPherson, S.R. & A. Robinson 2012. Field Guide to the Pitcher Plants of Borneo. Redfern Natural History Productions, Poole.
^ ^4.0 ^4.1 ^4.2 Clarke, C.M. 1997. Nepenthes of Borneo. Natural History Publications (Borneo), Kota Kinabalu.
^ Kurata, S. 1976. Nepenthes of Mount Kinabalu. Sabah National Parks Publications No. 2, Sabah National Parks Trustees, Kota Kinabalu.
^ Macfarlane, J.M. 1908. Ordo CLXXV bis. Nepenthaceæ. In: A. Engler Das Pflanzenreich IV, 111, Heft 36: 1–91.
^ Danser, B.H. 1928. The Nepenthaceae of the Netherlands Indies. Bulletin du Jardin Botanique de Buitenzorg, Série III, 9(3–4): 249–438.
^ Thong, J. 2006. Travels around North Borneo – Part 1.PDF Victorian Carnivorous Plant Society Journal 81: 12–17.

[Anonymous] 1887. Nepenthes culture.（页面存档备份，存于互联网档案馆） The Gardeners' Chronicle, series 3, 2(41): 442–443.
Beaman, J.H. & C. Anderson 2004. The Plants of Mount Kinabalu: 5. Dicotyledon Families Magnoliaceae to Winteraceae. Natural History Publications (Borneo), Kota Kinabalu.
Dixon, W.E. 1889. Nepenthes.（页面存档备份，存于互联网档案馆） The Gardeners' Chronicle, series 3, 6(144): 354.

扩展阅读

食虫植物照片搜寻引擎中哈里猪笼草的照片（页面存档备份，存于互联网档案馆）

维基共享资源上的相關多媒體資源：哈里猪笼草維基物種上的相關信息：哈里猪笼草

[Burbidge-6] 弗雷德里克·威廉·伯比奇对哈里猪笼草的叙述原文：^[2]

“ Apart from these I found an intermediate between N. villosa and N. Edwardsiana, also epiphytic on Casuarina. This is, I believe, unnamed ; if so, I should like it to be called Nepenthes Harryana. Now, if a dried pitcher of N. Edwardsiana be examined, the upper four-fifths of it will be seen to be membranous, the lower part leathery and hard ; in N. villosa nearly all is hard and leathery except about half-an-inch below the hardened rim of the urns ; in N. Harryana about one-third is hard, and two-thirds soft or membranous below the rim. The edge of the pitcher mouths in these three kinds is quite distinct from those of all others, as shown in my sketches.
[...]
Sir Joseph Hooker, in Linn. Trans., vol. xxii., suggested that N. villosa and N. Edwardsiana might be forms of the same species. This is not so, however, they are quite distinct in zone on the mountain, and in habit of growth also, and in colour. Then N. Harryana is a hybrid no doubt, which shows they have distinct sexual characters.
”

[1] von Arx, B., J. Schlauer & M. Groves 2001. CITES Carnivorous Plant Checklist.PDF The Cromwell Press, United Kingdom.

[Burbidge-2] 2.0 ^2.1 ^2.2 ^2.3 Burbidge, F.W. 1882. Notes on the new Nepenthes. （页面存档备份，存于互联网档案馆） The Gardeners' Chronicle, new series, 17(420): 56.

[3] McPherson, S.R. & A. Robinson 2012. Field Guide to the Pitcher Plants of Borneo. Redfern Natural History Productions, Poole.

[Clarke-4] 4.0 ^4.1 ^4.2 Clarke, C.M. 1997. Nepenthes of Borneo. Natural History Publications (Borneo), Kota Kinabalu.

[Kurata-5] Kurata, S. 1976. Nepenthes of Mount Kinabalu. Sabah National Parks Publications No. 2, Sabah National Parks Trustees, Kota Kinabalu.

[Macfarlane-7] Macfarlane, J.M. 1908. Ordo CLXXV bis. Nepenthaceæ. In: A. Engler Das Pflanzenreich IV, 111, Heft 36: 1–91.

[Danser-8] Danser, B.H. 1928. The Nepenthaceae of the Netherlands Indies. Bulletin du Jardin Botanique de Buitenzorg, Série III, 9(3–4): 249–438.

[Thong-9] Thong, J. 2006. Travels around North Borneo – Part 1.PDF Victorian Carnivorous Plant Society Journal 81: 12–17.

[1]

[2]

[3]

[4]

[5]

[注 1]

[6]

[7]

[8]

植物学史

相关物种

注释

参考文献

扩展阅读